Overrides are specific permissions designed to override a role in a specific context, allowing you to "tweak" your permissions as required.
In each role, you can choose to set the permission for a capability to one of four values:
Permissions at a "lower" context will generally override anything at a "higher" context (this applies to overrides and assigned roles). The exception is PROHIBIT which can not be overridden at lower levels.
Special exceptionsNote that the guest user account will generally be prevented from posting content even if granted the capability to do so.
See also Roles, Contexts, Assign Roles and Permissions.